Plant Physiology & Development

Stomatal Conductance & Gas Exchange

Stomata regulate the exchange of carbon dioxide and water vapor between the leaf and atmosphere. Their behavior responds to light, internal carbon dioxide, water status, vapor-pressure conditions, hormones, temperature, and other signals.

Key concepts

  • Carbon dioxide entry and water-vapor loss are coupled through stomatal pores
  • Stomatal conductance is not the same as photosynthetic rate
  • High atmospheric demand can increase transpiration pressure while also promoting stomatal restriction if plant water supply cannot keep pace
  • Leaf temperature and boundary-layer conditions change gas-exchange behavior
  • Root-zone stress can alter stomatal behavior before dramatic visual symptoms appear

What to measure or observe

  • Record air temperature, RH, leaf temperature, and light context together
  • Compare leaf posture and temperature across canopy zones
  • Observe recovery after irrigation or environmental transitions
  • Use gas-exchange instruments when quantitative stomatal conductance is required
  • Track root-zone moisture and EC alongside atmospheric measurements

Common mistakes

  • Treating a VPD number as a direct measurement of stomatal conductance
  • Assuming open stomata always mean optimal photosynthesis
  • Ignoring leaf temperature when interpreting atmospheric demand
  • Changing humidity without considering root water supply
  • Diagnosing stomatal behavior from leaf curl alone

Visuals this lesson still needs

  • Stomatal pore gas-exchange cutaway
  • CO2-in water-vapor-out diagram
  • Atmosphere-root coupling map
  • Leaf-temperature and boundary-layer graphic